WASHINGTON (AP) -- The Biden administration announced wide-ranging actions Wednesday meant to call out Russian influence in the upcoming U.S. presidential election, unsealing criminal charges against two employees of a Russian state-run media company and seizing internet domains used by the Kremlin to spread disinformation.

The measures represented a U.S. government effort at disrupting a persistent threat from Russia that American officials have long warned has the potential to sow discord and create confusion among voters. Washington has said that Russia remains the primary threat to elections even as the FBI investigates a hack by Iran of Donald Trump's campaign and an attempt breach of the Joe Biden-Kamala Harris campaign.

One criminal case accuses two employees of RT, a Russian-state-funded media organization that was forced by the Justice Department to register as a foreign agent, of covertly funding a Tennessee-based content creation company to publish nearly 2,000 videos containing Russian propaganda. The defendants, who remain at large, used fake identities and the company was unaware it was being used by Russia.

In the other action, officials announced the seizure of 32 internet domains that were used by the Kremlin to spread Russian propaganda and weaken global support for Ukraine.

Attorney General Merrick Garland said the actions relate to Russia's use of state media to enlist unwitting American influencers to spread propaganda and disinformation.

Intelligence agencies have previously charged that Russia was using disinformation to try to interfere in the election. The new steps show the depth of U.S. concerns and signal legal actions against those suspected of being involved.

"Today's announcement highlights the lengths some foreign governments go to undermine American democratic institutions," the State Department said. "But these foreign governments should also know that we will not tolerate foreign malign actors intentionally interfering and undermining free and fair elections."

In a speech last month, Deputy Attorney General Lisa Monaco said Russia remained the biggest threat to election integrity, accusing Russian President Vladimir Putin and "his proxies are using increasingly sophisticated techniques in their interference operations. They're targeting specific voter demographics and swing-state voters to in an effort to manipulate presidential and congressional election outcomes. They're intent on co-opting unwitting Americans on social media to push narratives advancing Russian interests."

Much of the concern around Russia centers on cyberattacks and disinformation campaigns designed to influence the November vote. The tactics include using state media like RT to advance anti-U.S. messages and content, as well as networks of fake websites and social media accounts that amplify the claims and inject them into American's online conversations. Typically, these networks seize on polarizing political topics such as immigration, crime or the war in Gaza.

In many cases, Americans may have no idea that the content they see online either originated or was amplified by the Kremlin.

"Russia is taking a whole of government approach to influence the election including the presidential race," an official from the Office of the Director of National Intelligence said this summer during a briefing. The official spoke on condition of anonymity under rules worked out with that office.

Groups linked to the Kremlin are increasingly hiring marketing and communications firms within Russia to outsource some of the work of creating digital propaganda while also covering their tracks, the officials said during the briefing with reporters.

Two such firms were the subject of new U.S. sanctions announced in March. Authorities say the two Russian companies created fake websites and social media profiles to spread Kremlin disinformation.

The ultimate goal, however, is to get Americans to spread Russian disinformation without questioning its origin. People are far more likely to trust and repost information that they believe is coming from a domestic source, officials said. Fake websites designed to mimic U.S. news outlets and AI-generated social media profiles are just two methods.

Messages left with the Russian Embassy were not immediately returned.